ulous) cones with exserted bracts, and male cones clustered in umbels, in these features more closely allied to the genus Keteleeria. Ecology The species are all adapted to (and are confined to) relatively moist, cool temperate areas with high rainfall, cool summers, and little or no water stress; they are also adapted to cope with heavy to very heavy winter snowfall and tolerate ice storms better than most other trees. Hemlock trees are more tolerant of heavy shade than other conifers; they are, however, more susceptible to drought. Threats The two eastern North American species, T. canadensis and T. caroliniana, are under serious threat by the sap-sucking insect Adelges tsugae (hemlock woolly adelgid). This adelgid, related to the aphids, was introduced accidentally from eastern Asia, where it is only a minor pest. Extensive mortality has occurred, particularly east of the Appalachian Mountains. The Asian species are resistant to this pest, and the two w estern American hemlocks are moderately resistant. In North America, hemlocks are also attacked by hemlock looper. Larger infected hemlocks have large, relatively high root systems that can bring other trees down if one falls. The foliage of young trees is often browsed by deer, and the seeds are eaten by finches and small rodents. Old trees are commonly attacked by various fungal disease and decay species, notably Heterobasidion annosum and Armillaria species, which rot the heartwood and eventually leave the tree liable to windthrow, and Rhizina undulata, which may kill groups of trees following minor grass fires that activate growth of the Rhizina spores. Uses The wood obtained from hemlocks is important in the timber industry, especially for use as wood pulp. Many species are used in horticulture, and numerous cultivars have been selected for use in gardens. The bark of the hemlock is also used in tanning leather. The needles of the heml