me species, differences in form between the sexes is quite marked. For example, in the African species Agrius convolvuli (the convolvulus or morning glory hawk moth), the antennae are thicker and wing markings more mottled in the male than in the female. Only males have both an undivided frenular hook and a retinaculum. Also, all male hawkmoths have a partial comb of hairs along with their antennae. Females call males to them with pheromones. The male may douse the female with a pheromone before mating. Behavior Some species fly only for short periods either around dusk or dawn, while other species only appear later in the evening and others around midnight, but such species may occasionally be seen feeding on flowers during the day. A few common species in Africa, such as the Oriental bee hawk (Cephonodes hylas virescens), Macroglossum hirundo, and Macroglossum trochilus, are diurnal. A number of species are known to be migratory, all in the
Sphingini and Macroglossinae, and specially in the genera Agrius, Cephonodes, Macroglossum, Hippotion and Theretra. Flight In studies with Manduca sexta, moths have dynamic flight sensory abilities due to their antennae. The antennae are vibrated in a plane so that when the body of the moth rotates during controlled aerial maneuvers, the antennae are subject to the inertial Coriolis forces that are linearly proportional to the angular velocity of the body. The Coriolis forces cause deflections of the antennae, which are detected by the Johnston's organ at the base of each antenna, with strong frequency responses at the beat frequency of the antennae (around 25 Hz) and at twice the beat frequency. The relative magnitude of the two frequency responses enables the moth to distingu
